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Introduction 1
1 Evolutionary Matters 10
2 Sociality, Media, and Variability 19
3 The Wisdom of Elders 29
4 Leaders 38
5 Teaching and Learning 50
6 Reproduction 60
7 Successful Subordinates 72
8 The Fall of Titans 80
9 Aging of Captive Alphas 93
10 Happy Families 100
11 Mothering - Good and Not So Good 111
12 Grandmothers 122
13 Sexy Seniors 134
14 Their Own Person 147
15 Adapting and Not Adapting 155
16 All Passion Spent 167
17 The Inevitable End 184
Notes 195
References 199
Index 217
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