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Preface ix
Theoretician's Laboratory 1
The structure of evolutionary theory 3
The utility of simple models 4
Why not just simulate? 8
A model of viability selection 11
Determining long-term consequences 16
Nongenetic replication 27
Animal Conflict 37
The Hawk-Dove game 38
Retaliation 46
Continuous stable strategies 52
Ownership, an asymmetry 55
Resource holding power 58
Sequential play 60
Altruism & Inclusive Fitness 71
The prisoner's dilemma 72
Positive assortment 76
Common descent and inclusive fitness 78
Rediscovering Hamilton's rule 82
Justifying Hamilton's rule 97
Using Hamilton's rule 99
Reciprocity 123
The Axelrod-Hamilton model 124
Mutants and mistakes 132
Partner choice 145
Indirect reciprocity 150
Reciprocity and collective action 156
Animal Communication 173
Costlysignaling theory 174
Cheap, honest signals 192
Signaling and altruism 201
Social learning 206
Selection among Groups 223
Three views of selection 225
Deriving the Price equation 228
Selection within and between groups 232
Dispersal 249
Sex Allocation 261
Fisher's theory of sex allocation 262
Reproductive value and Fisherian sex ratios 263
Using the Shaw-Mohler theorem 267
Biased sex ratios 271
Breaking the eigen barrier 281
Sexual Selection 295
Quantitative genetic models 298
Fisher's runaway process 305
Costly choice and sensory bias 309
Good genes and sexy sons 313
Appendixes 333
Facts about Derivatives 333
Facts about Random Variables 335
Calculating Binomial Expectations 337
Numerical Solution of the Kokko et al. Model 343
Solutions to Problems 349
Bibliography 393
Index 409
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